EZO-20406 Invertebrates, Algae & Fungi, no dissect


Credits 6.00

Teaching methodContact hours
Field practical9
Excursion (one day)3
Independent study0
Course coordinator(s)dr. SWS Gussekloo
Lecturer(s)dr. SWS Gussekloo
dr. ir. MJ Ketelaar
dr. ir. A Schouten
prof. dr. ir. FPM Govers
Examiner(s)dr. ir. MJ Ketelaar
dr. SWS Gussekloo

Language of instruction:


Assumed knowledge on:

Human and Animal Biology l and 2; Structure and Physiology of Plants; Growth, Development and Reproduction of Plants.

Continuation courses:

Vertebrate Structure and Function; Developmental Biology of Animals; Functional Zoology.


Note 1: Registration for dissection-free course is only possible after consultation with and via the course coordinator.
Note 2: The overall course (EZO 20306/20406) has a maximum number of participants. The deadline for registration is one week earlier than usual.
See Academic Year (http://www.wageningenur.nl/en/Education-Programmes/Current-Students/Agenda-Calendat-Adacemic-Year.htm) -> Registration for Courses.
Note 3: As postulated in the student charter a contribution has to be paid to cover the cost of living during the field course. 
Note 4: This course cannot be combined in an individual programme with EZO-22806 Marine life.

How have lower plants and invertebrate animals adapted in structure, function and behaviour to survive and reproduce in a wide spectrum of biotic and abiotic conditions? Such questions are discussed on phylogenetic and ontogenetic time scales. This provides insight in biodiversity, evolutionary theory and the social and economic importance of invertebrates and lower plants. Following lectures, histological, dissection and IT-supported practicals, all questions are finally integrated in a fieldcourse where animals and plants are studied in their natural habitat of the intertidal zone (Wimereux, France). There we can explore the adaptations and constraints of plant and invertebrate life, and their interactions in a natural community. You are also trained in doing research in a field work setting

Learning outcomes:

After successful completion of this course students are expected to be able to:
- ecognize and understand the differences in morphology and physiology between different groups (taxa/populations) of invertebrates, algae and fungi;
- understand the processes and mechanisms which shaped the organismal biological diversity: their adaptive radiation during the individual (ontogenetic) and the evolutionary (phylogenetic) development'
- master skills for your own research: observation, problem definition, analysis (including dissection), synthesis, time managment, discussion and presenting (oral, poster and written)'|- recognize and understand the diversity of lower plants and animals, in relation to abiotic (exposure, substrate, water content, salinity, temperature, oxygen content, pH) and biotic factors (community with other plants and animals) in the environment where they face selective forces;
(specific learning outcomes are given in each chapter of the syllabi and field guide)


In Wageningen:
- lectures and practicals on the subject of invertebrate biology, algae and fungi integrated with theoretical issues, and an interactive IT-supported practical programme on algae, fungi and lichens, including self-tests.
During the field work in Wimereux:
- general training in fieldwork and appropriate time management;
- during low-tide you will characterise (in biotic and physical sense) selected field locations;
- observed species are studied by teams of two students and analysed for taxonomy, structure, function, life style and biological role;
- eventually, 'simple' experiments exploring adaptation of animals and plants are designed and performed.
The results of a selection of the assignements are presented to the whole group, and/or written down in a lab journal.


The final grade includes four components, of which the weights will be communicated at the beginning of the course:
- computer test with closed questions on the subject of functional morphology of invertebrates;
- computer test with closed questions on the subject of general biology of invertebrates (e.g. evolution, adaptation, ecology);
- computer test with closed questions on the subject of the general biology of algae and fungi;
- field course performance. this grade is based on observation of laboratory/field participation (25%); (poster-) presentations (25%), and a lab/field journal including written assignments (50%).


- Hickman [et al.] Integrated Principles of Zoology;
- Raven [et al.] Biology of plants;
(both used in previous courses).
· Biologie van Evertebraten;
· Biologie van Algen en Schimmels;
· Introductie en Logboek Veldpracticum Evertebraten, Algen en Schimmels.
·  CD-ROM Biology of lower and higher plants;
·  Additional information in BrightSpace.